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Holocene coccoliths are predominant in the upper layers of Black Sea sediment cores, whereas reworked Eocene and Cretaceous coccoliths are a major part of the fine-grained carbonate material in lower levels of the Quaternary. Three stratigraphic units based on the distribution of these coccoliths reflect a rising worldwide sea level and an increase in salinity in the Black Sea at the time that marine circulation began through the Bosporus. The most dominant Holocene coccoliths of the Black Sea--Braarudosphaera bigelowi and Emiliania huxleyi--can thrive at the lower-than-normal marine salinity that now exists in the sea.
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